Lester the Jester was banished from his aphid colony. He travels throughout the insect world as a court jester trying to make a living with his comedy act. Secretly, Lester longs for the day when the aphid colony forgives him and welcomes him back with open arms.

Roach Recon are highly trained units of roaches. They are armed with a variety of weapons, especially handheld bombs, and they ride scramblers, light beetle tanks. Their main attack consists of blasting giant craters from underground and emerging with guns blazing.

Army Ants guard the ant hill. They man the bunkers, set the mines, and carry out suicide machine gun attacks. The ants ride stalkers, giant beetle tanks, and sky crawlers, in an never ending assualt against the insect world!

Slugs heads the criminal insect underworld. This ruthless kingpin controls all corrupt and illegal activity. Slugs carefully plans his escape from the ant dungeon.